Authors: Michael S. Batcher, Global Invasive Species Team, The Nature Conservancy
- Ligustrum spp. are deciduous, semi-evergreen, or evergreen shrubs and small trees in the Oleaceae (olive family). There are approximately 50 Ligustrum species that are native to Europe, North Africa, and Asia. Ligustrum spp. have been cultivated and developed into several horticultural varieties, and were introduced to North America as a common hedge in landscaping. Some Ligustrum spp. can grow to 5 m tall and have a stem diameter of 2.5-25 cm. Ligustrum spp. bark is whitish-tan to gray in color and smooth in texture. Slender twigs are straight, rounded or four-angled below the nodes, and gray-green in color. Winter buds are ovoid with two outer scales. Terminal buds are present.
- The simple leaves are elliptic to ovate in shape, oppositely arranged on slender twigs, often leathery and thick. The leaf margins are smooth and entire.
- Flowers have both male and female parts, and the corollas are white. The calyx is small, obconic or campanulate, and 4-toothed. Each flower has petals that are fused into a tube below with four separate lobes above. Flowers are borne on small panicles terminating the main axis and on short lateral branches. Bloom time is usually June-July.
- The fruit is a subglobose or ovoid drupe containing 1-4 seeds. Fruit clusters generally ripen during September and October and persist through the winter. Mature specimens can produce hundreds of fruit.
- Ecological Threat
- Ligustrum spp. can easily escape cultivation to invade adjacent areas and can form dense monospecific thickets. Several Ligustrum species have become common invaders of cultivated landscapes, disturbed areas and wildlands throughout the U.S.
Latin and Common Names:
Ligustrum amurense (Carr.): Amur privet
Ligustrum japonicum (Thun.): Japanese privet
Ligustrum lucidum (Ait.f.): Tree privet; glossy privet
Ligustrum obtusifolium (Sieb. and Zucc.): Blunt-leafed privet; border privet
Ligustrum ovalifolium (Hassk.): California privet; waxy-leaved privet
Ligustrum quihoui (Carr.): Wax-leaf privet
Ligustrum sinense (Lour.): Chinese privet
Ligustrum vulgare (L.): European privet, common privet
Ligustrum spp. are deciduous, semi-evergreen, or evergreen shrubs and small trees in the Oleaceae (olive family). There are approximately 50 Ligustrum species that are native to Europe, North Africa, and Asia. Ligustrum spp. have been cultivated and developed into several horticultural varieties, and were introduced to North America as a common hedge in landscaping. Ligustrum spp. can easily escape cultivation to invade adjacent areas and can form dense monospecific thickets.
Some Ligustrum spp. can grow to 5 m tall and have a stem diameter of 2.5-25 cm. Ligustrum spp. bark is whitish-tan to gray in color and smooth in texture. Slender twigs are straight, rounded or four-angled below the nodes, and gray-green in color. Winter buds are ovoid with two outer scales. Terminal buds are present. Leaves are elliptic to ovate in shape, oppositely arranged on slender twigs, often leathery and thick. Flowers have both male and female parts, and the corollas are white. The calyx is small, obconic or campanulate, and 4-toothed. Each flower has petals that are fused into a tube below with four separate lobes above. Flowers are borne on small panicles terminating the main axis and on short lateral branches. Bloom time is usually June-July. The fruit is a subglobose or ovoid drupe containing 1-4 seeds. Fruit clusters generally ripen during September and October and persist through the winter. Mature specimens can produce hundreds of fruit.
- Ligustrum amurense grows to 5 m. Leaves are elliptic to oblong or oblong-ovate, 2.5-6 cm long, acute or obtuse,
rounded or broad-cuneate at base; ciliolate, sometimes lustrous above, and smooth except on the midrib below. Petioles are 2-4 mm long, pubescent. Panicles are 3-5 cm long and pubescent. The calyx is glabrous or slightly pubescent. The corollas (from base of tube to tip of lobe) are 7-9 mm long, with the tube far longer than the lobes.
- Ligustrum japonicum generally grows to 3 m, rarely to 6 m. Leaves are broad-ovate to ovate-oblong, 4-10 cm long,
obtusely short-acuminate or acute to obtuse, rounded at the base with reddish margins and midrib and with 4-5 pairs of indistinct veins. Petioles are 6-12 mm long. Panicles are 6-15 cm long. Flowers are short-stalked with the corolla tube longer than the calyx. Stamens are slightly longer than the corolla lobes.
- Ligustrum lucidum grows as a large shrub or medium-sized tree, to 10 m high, with spreading branches. Leaves are
ovate to ovate-lanceolate, 8-12 cm long, acuminate or acute, usually broad-cuneate with 6-8 veins, usually distinct above and beneath. Petioles are 1-2 cm long. Panicles are 12-20 cm long and nearly as wide. Flowers are subsessile. The corolla tube is as long as the calyx. Stamens are as long as the corolla lobes. Fruits are oblong, 1 cm long, bluish or purplish-black.
- Ligustrum obtusifolium grows to 3 m with spreading or arching branches. Leaves are elliptic to oblong or oblongobovate,
2-6 cm long. Leaves are acute or obtuse, cuneate or broad-cuneate, glabrous above, pubescent below (or occasionally only on midrib). Petioles are 1-4 mm long, pubescent. Panicles are 2-3.5 cm long, nodding. Corollas are 8-10 mm long with anthers nearly as long as the corolla lobes. Fruits are subglobose, black and slightly bloomy (glaucous).
- Ligustrum ovalifolium grows to 5 m. Leaves are elliptic-ovate to elliptic-oblong, 3-6 cm long, acute, broad-cuneate,
dark lustrous green above, yellowish green below. Petioles are 3-4 mm long. Flowers are creamy-white with an unpleasant scent, subsessile in panicles 5-10 cm long. Corollas are 8 mm long with anthers as long as lobes. Fruits are 5-7 mm across, black.
- Ligustrum quihoui grows to 2 m with spreading, rigid branches. Leaves are elliptic to elliptic-oblong or obovate to
obovate-oblong, 2-5 cm long, obtuse, sometimes emarginate, glabrous, subcoreaceous. Petioles are 1-3 mm long, puberulous. Flowers are sessile, in small clusters on long spikes collected into 10-20 cm long panicles. Corolla tubes are as long as the lobes with anthers exceeding the lobes. Flowers appear in late summer.
- Ligustrum sinense is a shrub or small tree to 7 m. Leaves are elliptic to elliptic-oblong, 3-7 cm long, acuminate,
acute to obtuse, dull green above, pubescent on the midrib below. Petioles are 6-15 mm long. Flowers are small, distinctly stalked, on panicles 10-16 cm long. Fruits are dull black. Ligustrum vulgare grows to 5 m with spreading branches. Leaves are oblong-ovate to lanceoloate, 3-6 cm long, obtuse to acute, glabrous. Petioles are 3-10 mm long. Flowers are pediceled in dense puberulous panicles, 3-6 cm long. Anthers exceed the corolla tube. Fruits are subglobose or ovoid, 6-8 mm long, black and lustrous.
Several Ligustrum species have become common invaders of cultivated landscapes, disturbed areas and wildlands throughout the U.S. Ligustrum amurense is found in many eastern and some south-central states. L. japonicum is found in the Southeast and in Puerto Rico. Ligustrum lucidum is present from Maryland south and west to Texas. Ligustrum sinense and Ligustrum obtusifolium are found throughout the eastern and central U.S. Ligustrum ovalifolium is common in California and in parts of the central and eastern U.S. Ligustrum quihoui is seen in the southeast. Ligustrum vulgare is widely naturalized throughout much of the U.S. and southern Canada.
Ligustrum spp. may invade natural areas such as floodplain forests and woodlands. They may displace shrubs in regenerating communities and remain persistent in these areas. Ligustrum spp. can form dense thickets that outcompete many kinds of native vegetation.
In North America, Ligustrum spp. are seen along roadsides, in old fields and in other disturbed habitats and in a variety of undisturbed natural areas, including bogs, wetlands, floodplains, old fields, calcareous glades and barrens, and mesic hardwood forests.
Ligustrum spp. control methods include mowing and cutting, seedling removal, herbicide application, and burning. Mowing and cutting are appropriate for small initial populations or environmentally sensitive areas where herbicides cannot be used. Stems should be cut at least once per growing season as close to ground level as possible. Repeated mowing or cutting will control the spread of Ligustrum spp., but may not eradicate it. Ligustrum spp. can also be effectively controlled by manual removal of young seedlings. Herbicide control measures include foliar spraying in late autumn or early spring with glyphosate, triclopyr, or metsulfuron; cut stump applications using glyphosate or triclopyr; and basal bark applications of triclopyr. Some reports indicate that burning top-kills L. vulgare and L. sinense and, if repeated, can eliminate them over time.
Ligustrum spp. are native to east Asia, Europe and North Africa: Ligustrum amurense is native to north China; L. japonicum to Korea and Japan; L. lucidum to China, Korea and Japan; L. obtusifolium to Japan; L. ovalifolium to Japan; L. sinense to China; and L. vulgare to the Mediterranean region. Reported occurrences of the different Ligustrum spp. in North America include:
- Ligustrum amurense: Arkansas, Kentucky, Maine, Maryland, Massachusetts, New Jersey, New York, North Carolina, Pennsylvania, South Carolina, Tennessee, Texas, and Virginia.
- Ligustrum japonicum: Alabama, Florida, Georgia, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Texas, Virginia, and Puerto Rico.
- Ligustrum lucidum: Alabama, Florida, Georgia, Louisiana, Maryland, Mississippi, North Carolina, and Texas.
- Ligustrum obtusifolium: Connecticut, District of Columbia, Illinois, Indiana, Kentucky, Maryland, Massachusetts, Michigan, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, Tennessee, Utah, Vermont, and Virginia.
- Ligustrum ovalifolium: California, Connecticut, Delaware, District of Columbia, Florida, Kentucky, Louisiana, Maryland, Massachusetts, Michigan, Missouri, New Jersey, North Carolina, Ontario, Pennsylvania, Texas, Vermont, Virginia, and Puerto Rico.
- Ligustrum quihoui: Louisiana, North Carolina, Texas, and Virginia.
- Ligustrum sinense: Alabama, Arkansas, Connecticut, Florida, Georgia, Iowa, Kentucky, Louisiana, Maryland, Massachusetts, Mississippi, Missouri, New Jersey, North Carolina, Oklahoma, Rhode Island, South Carolina, Tennessee, Texas, and Virginia.
- Ligustrum vulgare has the broadest range of the invasive Ligustrum species established in North America. It has been documented in: Alabama, Arkansas, British Columbia, Connecticut, Delaware, District of Columbia, Florida, Georgia, Great Smoky Mountain National Park, Illinois, Indiana, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Missouri, New Hampshire, New Jersey, New York, Newfoundland Island (Newfoundland), North Carolina, Nova Scotia, Ohio, Ontario, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Utah, Vermont, Virginia, West Virginia, and Wisconsin.
Impacts and Threats
Ligustrum spp. can form dense thickets that outcompete native vegetation. The privets can invade natural areas such as floodplain forests, woodlands, and disturbed agricultural fields. They generally expand along fence-rows, windbreaks and roadsides. In New Zealand, L. sinense may displace the shrub layer and marginal shrubs of alluvial forests, and remain persistent in these areas. L. lucidum replaces midcanopy trees in forests and may completely dominate an area of forest or forest fragments if not controlled. Ligustrum japonicum and L. sinense invade woodlands in the eastern and southeastern U.S.. Forest gaps can also become invaded since birds often disperse Ligustrum seeds.
In North America, Ligustrum spp. often grow along roadsides, in old fields and in other disturbed habitats and in a variety of undisturbed natural areas. Examples of Ligustrum invasions include:
1) Ligustrum obtusifolium was found invading an old field succession site in Illinois. The field had an average of more than 6,082 plants per ha (2.5 acres).
2) Ligustrum sinense has been reported in bogs, an oak-hickory-pine forest, a longleaf pine-turkey oak forest, and mesic hardwood forests in Alabama. In Arkansas, L. sinense has been reported in virtually all non-xeric habitats. In Georgia, L. sinense has been reported in floodplain/wetland habitats, and in North Carolina, in woodland edges.
3) Ligustrum vulgare has been recorded in bottomlands and mesic and riparian forests in Arkansas. In Ohio, L. vulgare is found in old fields, primary woodlands, and closed canopy forests. In Tennessee, the species has been recorded in calcareous glades and barrens and in deciduous cove forests.
4) In New Zealand, L. sinense is found in alluvial forest remnants, waste places, shrublands, and open stream systems, particularly in coastal areas. Ligustrum sinense is widespread and common, especially near towns. It is a common farm hedging plant. L. lucidum is found in forests (lowland and coastal), forest fragments, shrublands, along roadsides, in farm hedges, wastelands, and domestic gardens.
Ecology and Biology
Ligustrum spp. are perennial shrubs that grow readily from seed or from root and stump sprouts. They can escape from cultivation when the fruits are consumed by wildlife, particularly birds, which often excrete the seeds unharmed at distant locations where they may germinate and become established. Germination rates have been variously reported as low as 5%-27% and as high as 77%. Unlike most woody species, experimental defoliation did not result in reduced percentages of flowers producing fruits, decreased seed number, or decreased seed quality.
Ligustrum spp. leaves are high in phenolic compounds that defend against herbivores, especially insects. These work by inhibiting digestive enzymes and proteins. Despite this, L. sinense has been identified as an important forage plant for deer in the southeastern U.S..
Ligustrum vulgare grows well in high light, low nutrient soils, but will tolerate lower light levels if nutrients are increased.
Potential for Restoration of Invaded Sites
In North America, Ligustrum spp. have no important pests or predators. The various species are widespread and occasionally locally abundant. Manual and mechanical, environmental/cultural, and chemical methods are all useful in varying degrees in controlling Ligustrum spp. Fire management may be useful in some cases where the density of Ligustrum spp. is low and sufficient fuels available. Restoration potential is likely to be lowest where Ligustrum spp. occur in high densities and there is a high likelihood of continued dispersal of seeds into the restoration area. Ligustrum spp. have a high degree of reproductive vigor, a wide range of adaptability, and, in its present settings, few pests and predators. Ligustrum spp. produce large numbers of viable seed that are readily dispersed by birds and germinate at high rates in a wide range of conditions.
The potential for large-scale restoration of unmanaged natural areas or wildlands infested with Ligustrum spp. is probably low. Restoration potential for managed natural areas or wildlands infested Ligustrum spp. is probably moderate. If attacked during the early stages of colonization, the potential for successful management is high.
Mowing and cutting are appropriate for small populations or environmentally sensitive areas where herbicides cannot be used. Stems should be cut at least once per growing season as close to ground level as possible. Repeated mowing or cutting will control the spread of Ligustrum spp., but may not eradicate it. Managers of The Nature Conservancy preserves in Ohio reported eradication of L. vulgare after two cutting treatments.
Ligustrum spp. can be effectively controlled by the manual removal of young seedlings. Plants should be pulled as soon as they are large enough to grasp but before they produce seeds. Seedlings are best pulled after a rain when the soil is loose. Larger stems (up to 6 cm in diameter) can be removed using a weed wrench or similar uprooting tools. The entire root must be removed since broken fragments may resprout.
Ligustrum spp. have no known biological controls, although a few pathogens are known to attack them in North America. Cercospora adusta, C. lilacis, and Pseudocercospora lugustri are fungal leaf spots that affect L. vulgare and L. amurense. Nectriella pironi creates galls on L. sinense, L. lucidum and L. quihoui. Pseudomas syringae impacts members of the olive family including L. amurense. Agrobacterium tumefaciens, Ganoderma lucidum and Glomerella cingulata affect L. vulgare.
Foliar Spray Method: This method may be effective for large thickets of Ligustrum spp. where risk to nontarget species is minimal. Air temperatures should be above 17°C (63°F) to ensure that herbicides are absorbed. The ideal time to treat is while plants are in leaf in late autumn or early spring but when many native species are dormant.
Glyphosate (brand name Roundup and others): A number of concentrations have been used successfully. The Tennessee Exotic Pest Plants Council (1996) suggests a 2% solution of glyphosate and water plus a 0.5% non-ionic surfactant to thoroughly wet all leaves. The New Zealand Weeds Web Site (1999) recommends, for a handgun sprayer, 1 liter Roundup and 100 mls of a surfactant per 100 liters of water (1% solution); for a backpack sprayer, the recommendation is 100 ml Roundup and 20 mls of a surfactant per 10 liters of water. (Roundup is a non-selective herbicide.)
Triclopyr (brand name Garlon, Pathfinder II and others): The Tennessee Exotic Pest Plants Council (1996) suggests a 2% solution of triclopyr and water plus a 0.5% non-ionic surfactant, sprayed to thoroughly wet all leaves. Use a low pressure and coarse spray pattern to reduce spray-drift damage to non-target species. (Triclopyr is a selective herbicide for broadleaf species only.)
Metsulfuron (brand name Escort and others): The New Zealand Weeds Web Site (1999) recommends, for a handgun sprayer, 35 g metsulfuron and 100 mls of a surfactant per 100 liters of water; for a backpack sprayer, the recommendation is 5 g metsulfuron and 10 mls of a surfactant per 10 liters of water. Metsulfuron methyl was identified as the most cost-effective herbicide in an experimental treatment comparing metsulfuron methyl, triclopyr ester and 2,4-D. (Metsulfuron is a selective herbicide active upon broadleaf and some annual grass species.)
Cut Stump Method: This control method should be considered when treating individual shrubs or where the presence of desirable species precludes foliar application. The Tennessee Exotic Pest Plants Council (1996) recommends this treatment only as long as the ground is not frozen, but other researchers have found it effective on Rhamnus spp. in frozen ground. Immediately after cutting stems at or near ground level, apply a 25% solution of glyphosate and water or triclopyr and water to the cut stump, being careful to cover the entire surface. Effectiveness of the herbicide is increased if holes are cut in the top of the freshly felled stump, to hold the herbicide in for better absorption by plant.
Basal Bark Method: Apply a mixture of 25% triclopyr and 75% horticultural oil to the basal parts of the shrub to a height of 30-38 cm (12-15 in) from the ground. Thorough wetting is necessary for good control; spray until run-off is noticeable at the ground line. Like the cut stump application, this method may be effective throughout the year, if Ligustrum spp. responds similarly to Rhamnus spp.. In New Zealand, researchers have killed standing Ligustrum trees by drilling downward-sloping 20 mm wide holes 5 cm into the trunk at no greater than 5 cm spacing around the trunk, and filling the holes with a stump paint-herbicide mix.
Faulkner et al. (1989) reported that in experimental trials of prescribed burning, there was no significant difference in the abundance of L. sinense in burned vs. unburned plots. Ligustrum litter has a low flammability and fires did not carry well in these treatments.
The Nature Conservancy land managers in Alabama reported that burning top-kills L. vulgare and L. sinense and eliminates them over time, and that burning is effective at controlling L. sinense if done annually with low fuel moisture and high Keetch-Byram Drought Index.
Examples of Ligustrum spp. Control on TNC Preserves
Ligustrum spp. have been reported as problems weeds on TNC preserves in Alabama, Arkansas, Louisiana, Georgia, Florida, Mississippi, Tennessee, North Carolina, and in Ohio.
In Alabama and in Florida, Carlen Emanuel and Greg Seamon, respectively, reported that annual burning was effective in controlling L. sinense. Furthermore, cutting is also effective if done when conditions are dry. George Ramseur Jr. in Mississippi found that a combination of pulling and burning provided good control of L. sinense.
Richard Martin reports that L. sinense is one of the worst weeds on Louisiana preserves, and has found that the application of Garlon 4 (triclopyr) has produced excellent control results, but RoundUp (glyphosate) did not provide good results. In North Carolina, however, Robert Merriam found that RoundUp was useful in controlling large infested areas of L. sinense. Additionally, cutting was very effective if coupled with the use of Arsenal (imazapyr) on cut stumps. Rates of herbicide application should follow those recommended by the manufacturer. Rates that have been applied successfully for control of Ligustrum are described above.
In Arkansas, Scott Simon reports that burning only top-kills L. vulgare and L. sinense, but will eventually eliminate the plants over time if burns are repeated. Burning is not effective however, in moist bottomland areas.
Ligustrum vulgare was successfully controlled in central Ohio preserves. Ross Lebold reported that the cut-stump method, using RoundUp (glyphosate) was effective, and that repeated cutting also seemed effective. In Tennessee, L. vulgare was partially controlled by cutting, and Gabby Call reports that the use of goats to control privet works well. The goats however, must able to reach and destroy adult privet plants.
- Gabby Call
The Nature Conservancy
50 Vantage Way, Suite 250
Nashville, TN 37228
- Carlen M. Emanuel
Alabama Natural Heritage Program
1500 East Fairview Avenue
Montgomery, AL 36106-2148
- Carol Helton
Conservation Program Coordinator
Atlanta Botanical Garden
P.O. Box 77246, Atlanta, GA 30357
- Ross Lebold
The Nature Conservancy
6375 Riverside Drive Street 50
Dublin, OH 43017
- Richard Martin
The Nature Conservancy
P.O. Box 4125
Baton Rouge, LA 70821
- Sam Pearsall
The Nature Conservancy
North Carolina Field Office
4011 University Dr.
Durham, NC 27707
- George Ramseur, Jr.
1709 Government St.
Ocean Springs, MS 39564
- Greg Seamon
P.O. Box 393
Bristol, FL 32321
- Scott Simon
Arkansas Field Office
601 N. University Ave.
Little Rock, AR 72205
- S. Lee Stone
Acting Chief Biologist
Natural Resources Division
Austin Parks and Recreation Department
301 Nature Center Drive
Austin, TX 78746
In natural areas management, monitoring programs will likely follow changes in abundance of Ligustrum spp. AND changes in abundance of desirable native species or changes in community attributes that are the targets of management. Such programs should have explicit objectives that can be measured and that are meaningful from both a biological and management standpoint. These objectives may vary depending on the abundance of Ligustrum spp. and other invasive plants. For instance, the objective of managing a forest with 40% cover of Ligustrum spp. may be to reduce Ligustrum cover to 20%. On the other hand, on a site with 10% an appropriate management objective might be to prevent an increase of more than 10% of total cover (20% total). In addition, increasing regeneration of native species may be an important objective. Monitoring the status of other conservation targets such as invertebrates dependent on specific nectar sources or plant species that are conservation targets may be more important than tracking invasive plant species abundance. In general, the objectives of monitoring should track those of management.
In terms of effort (number of plots established and monitored), transects or long linear plots are more effective in providing the statistical power to necessary to detect changes than square, broadly rectangular, circular or other regularly shaped quadrats. Analyses of plant species composition and abundance can be simplified by (1) collecting data on abundance of dominant species; (2) collecting data on all species and pooling data on less abundant species; and (3) pooling data on species by placing them in guilds (e.g. invasive grasses, invasive legumes, native grasses, etc.).
While generally a research technique, measuring change, or lack thereof, in control (unmanaged) areas can be an effective way of assuring that changes observed in treated areas actually result from the treatmentand not from other factors such as fire, rainfall, etc. In forest communities that are in early successional stages or recently disturbed, declines in abundance of the Ligustrum spp. may occur with time without management.
Additional research is needed on more efficient control methods, especially where cutting is used. Standard tools such as weed whackers, brush hogs, and other equipment are not designed for cutting this species or for use in the kinds of habitat which Ligustrum species often invade.
- Rehder, A. 1977. Manual of cultivated trees and shrubs hardy in North America. Macmillan, NY.
- Haragan, P.D. 1996. Ligustrum vulgare, L. sinense, L. japonicum. pp.58-58 in Randall, J. and J. Marinelli (eds.). Invasive Plants: Weeds of the Global Garden. 1997. Brooklyn Botanic Garden, NY.
- ↑ New Zealand Weeds Web Site:www.boprc.govt.nz/www/green/weedsindx.html (Accessed 1999). 3.0 3.1 3.2 3.3 3.4 3.5
- Stone, S.L. 1997. Privet removed from Austin Nature Preserves (Texas). Restoration and Management Notes. 15(1): 93.
- ↑ Faulkner, J.L.; E.E.C. Clebsch. 1989. Use of prescribed burning for managing natural and historic resources in Chickamauga and Chattanooga National Park, USA. Environmental Management. 13(5): 603-312. 5.0 5.1
- ↑ Tennesee Exotic Pest Plants Council. 1996. Invasive Exotic Pest Plants of Tennessee, http://www.tneppc.org/. 6.0 6.1 6.2 6.3 6.4 6.5 6.6 6.7
- ↑ Randall, J.M. and B.A. Meyers-Rice. unpublished. 1998 Weed Survey of The Nature Conservancy’s land managers. Documents on file at TNC Wildland Invasive Species Program, Davis, CA 7.0 7.1 7.2 7.3
- Schopmeyer, C.S. 1974. Seeds of Woody Plants in the United States. Agriculture Handbook No. 450. Forest Service, USDA, Washington, D.C.
- Obeso, J.R. and P.J. Grubb. 1993. Fruit maturation in the shrub Ligustrum vulgare (Oleacea): lack of defoliation effects. Oikos 68: 309-316.
- Konno, K.; H. Yasui; C. Hirayama; H. Shinbo. 1998. Glycine protects against strong protein denaturing activity of oleuropein, a phenolic compound in privet leaves. Journal of Chemical Ecology. 24(4): 735-751.
- Stromayer, K.A.; R.J. Warren; A.S. Johnson; P.E. Hale; C.L. Rogers; C.L. Tucker. 1998. Chinese privet and the feeding ecology of white-tailed deer: the role of an exotic plant. Journal of Wildlife Management,61(4): 1321-1329.
- Grubb, P.J.; W.G. Lee; J. Kollmann; J.B. Wilson. 1996. Interaction of irradiance and soil nutrient supply on growth of seedlings of ten European tall-shrub species and Fagus sylvatica. Journal of Ecology. 84: 827-840.
- Sinclair, W.A.; H.H. Lyon; and W.T. Johnson. 1987. Diseases of trees and shrubs. Cornell University Press,Ithaca, NY.
- Madden, J.E. and J.T. Swarbrick. 1990. Chemical control of Ligustrum lucidum. Plant Protection Quarterly.5(4): 145-147.
- ↑ Reinartz, J.A. 1997. Controlling Glossy Buckthorn (Rhamnus frangula L.) with winter herbicide treatments of cut stumps. Natural Areas Journal. 17(1). 38-41. 15.0 15.1
- Buchanan, R.A. 1989. Pied currawongs (Strepera graculina): their diet and role in weed dispersal in suburban Sydney, New South Wales. Proceedings of the Linnean Society of New South Wales, 111(4):241-255.
- Radford, A.E., H.A. Ahles and C.R. Bell. Manual of the Vascular Flora of the Carolinas. 1964.
- Taylor, G. and S. Killiffer. 1996. Exotic plant species profile: Common privet, Ligustrum spp. TN-EPPC News.
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